Lupe, an impala with a flair for mechanics, Flapjack, a sweet young octopus with vision problems, and the other Lowriders are concerned about the pollution in their town, and the greedy Upscale Business Association's plans to tear down local businesses--but first they must rescue the migrating monarchs from a wildfire caused by drought and climate change.

Lupe, Flapjack, and Elirio customize their car into a low rider for the Universal Car Competition to win the cash prize that will enable them to buy their own garage.